1 Program in Molecular Medicine, University of Massachusetts Medical School, Worcester, MA, USA, 2 Broad Institute of Harvard and MIT, Cambridge, MA, USA, 3 Program in Bioinformatics and Integrative Biology, University of Massachusetts Medical School, Worcester, MA, USA, 4 Department of Neurobiology and Brudnick Neuropsychiatric Research Institute, University of Massachusetts Medical School, Worcester, MA, USA, 5 Department of Psychology, Stanford University, Palo Alto, CA, USA, 6 Wu Tsai Neurosciences Institute, Stanford University, Palo Alto, CA, USA, 7 Sanderson Center for Optical Imaging and Department of Microbiology and Physiological Systems, University of Massachusetts Medical School, Worcester, MA, USA, 8 Department of Stem Cell and Regenerative Biology, Harvard University, Cambridge, MA, USA, 9 Department of Psychiatry, Massachusetts General Hospital, Boston, MA, USA