In this paper we explore territorial behavior under the ecological conditions that fostered conflict in our evolutionary past. We develop an economic model that isolates how resource skew and variance affect territorial ranges, as well as how they interact with unequal appropriation abilities. Testing the predictions in a controlled laboratory experiment, we find that observed behavior clusters around the equilibrium predictions and all comparative statics have the predicted sign. We also find that equally strong appropriators choose more exclusive and less overlapping ranges than predicted with symmetric resources, but weaker appropriators choose more engulfed ranges than predicted with skewed resources.